I read that a few databases can be used in-memory but can’t think of reason why someone would want to use this feature. I always use a database to persist data and memory caches for fast access.
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Cache is also a kind of database, like a file system is. ‘Memory cache’ is just a specific application of an in-memory database and some in-memory databases are specialized as memory caches.
Other uses of in-memory databases have already been included in other answers, but let me enumerate the uses too: